The Ane have been unwilling to discuss exactly what the Speakers do except to state they have no governmental authority. If pressed they will say they have a function in the cross dimensional interface with the All. Not something that can be described in words. There is a First Speaker, Zenderban. He became First Speaker some time in the 19th century. Speakers are reluctant to speak with non telepaths. Apparently they are constantly in full contact with the All.
Because the speakers of the All are so connected to the All that are spacier than the average Ane by far. They find it difficult to communicate with non-telepaths without bulldozing you with the All itself.
Most speakers are the more typical introverted Ane and are never even seen by those not on Ane worlds. The numbers of speakers is unknown.